A few more case studies:

     19) Blue whale.

          We haven't looked at any purely aquatic species yet.  Whales are also a "big
          attention getter" for conservation (I mean seriously - did anyone hear of species
          like the dark-rumped petrel before taking this class?).

          - This species emphasizes the need for international cooperation for it's
          protection.

          - It also shows that some environmental problems are global.

          Natural history:

          As everyone knows, this is the largest animal to exist that we know about:

               - Size at sexual maturity is 22.5 meters in males and 25 meters in females
               (antarctic specimens slightly larger)

               - record is 33.58 m female (110 feet) for length, and 190,000 kg (419,000
               lbs.) female for weight.

          Baleen whales - feed on almost solely on crustaceans of the family Euphausiidae
          (generally less than 5 cm (2 inches) long).  Daily consumption has been estimated
          as high as 40 million individuals (3,600 kg).  

               - Feeds only during summer.  Rest of the year doesn't feed at all but lives
               off stored fat.

          Low  birth rate, females give birth every 2 - 3 years, almost always to a single
          young.  Gestation period is about 9 « months.

               - at birth the young is about 7 m, and 2,000 kg.

               - sexual maturity is at 5 - 10 years.

          Life span estimates go up to 110 years, though generally thought to be somewhere
          around 60 (?).  

          Some of this life-history information is rather conjectural.

          - Initially difficult to hunt, automation made this much easier.  It is estimated that
          up to 350,000 individuals were killed during the 20th century.  (1930/31 records
          show 29,410).

               - Numerous efforts launched to protect the whale since the 1930's, but all
               failed due to lack of compliance.  By the 1960's the whale was nearing
               extinction.

               - One problem that emerged is that populations may be too low to be
               viable.  For example, in 1986 it was estimated that 1,300 to 2,000 whales
               were left in the souther hemisphere (from a population of 150,000 -
               210,000).  In 1989 the estimates had dropped to 500 whales.  Antarctic
               populations do not show any recovery.

               - There does seem to be an increase in other areas (Iceland, California). 
               But total numbers are hard to gauge.  There are also conflicting reports
               about whales around Iceland.

               -  The populations don't mix much since they migrate at different times
               (i.e., southern hemisphere populations move north when northern
               hemisphere populations move north).

               - One estimate (2000, World Wildlife, puts the total population at under
               5,000, though this excludes the "Pygmy blue whale" subspecies (how's
               that for an oxymoron!)

          - New threats are coming from two possible sources:

               - Humans are now harvesting krill in increasing numbers for food.

               - There is some speculation that global warming may adversely affect krill
               populations.

          - The main cause of the problem (whale hunting) has been eliminated.  There are a
          few violation (e.g., Japan's infamous "research" hunting).  It is too early to know
          if the species will make a come-back.

               - Protection did not start until the mid 1960's.  All whale hunting (not just
               blue whales) was banned after 1984/85.  

               - The marine mammal protection act (1972) was pivotal in this.  It
               prohibited any taking of whale products by anyone subject to U.S.
               jurisdiction.

          - Other whale populations are obviously also endangered, though you'll have to
          look up the details yourself.

          - An unrelated anecdote about killer whales:

               - They are known to prey on other whales.  For example, marine biologists
               have observed a pod of killer whales decimating a small group of sperm
               whales (essentially taking by taking big chunks out of the whales until the
               bled to death).  There are sometimes purely natural dangers to animals. 
               (Killer whales definitely deserve the name!).

     20) Cheetahs (a personal favorite).

          - Illustrates a lot of misconceptions.

          - Fastest land animal, as most folks know.  Can reach speeds up to 70 mph (for
          very short distances!).  Incidentally, number 2 on the speed list is thought to be the
          pronghorn.

          Range used to go through most of Africa, through the Middle East, spreading into
          Iran, Pakistan and India.  

               There are "field" guides from the 1800's put out by the British that
               describe cheetahs in India.

          Prehistoric cheetahs existed in North America.  These were rather larger than
          present day forms.

          Until recently, was thought to be extinct outside of Africa.  Consistent reports of
          sightings indicate that some may still exist in Iran/Pakistan/Afghanistan.  (Anyone
          want to go find out?  This does illustrate some problems about getting
          information).

               There are also reports that the cheetah may be able to cope with extremely
               low population densities.  This might explain it's persistence without
               being regularly seen.

          Natural history:

               Several good studies have been done on the cheetah, mostly in East Africa,
               though some from other areas (e.g., Shaller, Caro, Kruuk, Burney).

               Basic social structure seems to be solitary, though related males may
               sometimes stay in small groups (2 - 3).

               Litter size is variable, though up to 6 is not uncommon.  Mortality can be
               very high in some areas (some estimates are up to 90% in the Serengeti).

               Young stay with mother for just about 18 months.

               Hunt mostly small gazelles.  But larger prey may be taken occasionally,
               particularly by cheetahs living in groups (e.g.,  zebra or wildebeest).  In
               areas without gazelles, are opportunists.  A study in Namibia showed
               cheetahs living mostly off rabbits.

               Are adapted for running down prey with a sudden burst of speed.  Often
               the prey is tripped with a leg, and then suffocated by clamping down on
               the throat.

               Prey is often stolen by other predators.  Cheetahs are not powerful fighters,
               so hyenas, lions, and even leopards will steal their prey.  Cheetahs have
               adapted to eat their meals quickly.  

               No natural "predators", though lions will often kill cheetahs, contributing
               substantially to adult mortality.

               Tame down readily.  If the breeding problem (see below) could be solved,
               they might even make decent pets, though that's a whole different issue.

                    - were accorded privileged status in many cultures

                    - were used for "hunting".  Similar to falconeering - cheetahs were
                    taken on a chariot until this got close to a herd of gazelle.  A hood
                    was then removed from the cheetah, and it was released.  The
                    cheetah would run down the gazelle.  

                         - this was done mostly for sport, not for meat.

                         - records show that during the 16th century one Indian
                         mogul kept over 9,000 cheetahs during his life time (but
                         only one birth was recorded).

          Conservation status:

               Estimates are hard to come by, but indicate that in 1900 more than
               100,000 cheetahs existed throughout Africa and Asia.

               Today the numbers have dropped to somewhere between 9,000 and
               12,000, though exact numbers are difficult to come by.  Asian population
               (in Iran and Afghanistan is estimated at just over 200).

                    - Concentrated in East and South Africa.  In some parts they still
                    exist at "normal" population levels.

               Main threats are from:

                    - encroaching human populations and hunting (which persists in
                    some areas despite protection as an endangered species - they are
                    accused, for instance, of taking live-stock in Namibia).    

                    - other predators, which will kill cubs and even adults.  See
                    estimated mortality figure above.

               As such, it turns out that cheetahs sometimes do best in areas just outside
               reserves where larger predators are absent.

                    - It is probably too early to make recommendations, but one
                    possibility for helping cheetahs might be to set up areas where
                    other large predators are excluded (possibly difficult since at least
                    lions are also listed as threatened).

               Captive breeding has simply not worked.  Despite numerous efforts, only
               sporadic success has been reported.  Possible reasons include:

                    - high percentage of defective sperm (but they seem to manage in
                    the wild!)

                    - an incomplete knowledge (even after all the studies) of breeding
                    biology.  

                         For example, one theory suggests that cheetahs do not
                         breed unless there is sufficient prey around.  In zoos, they
                         do not "see" their prey, so breeding is suppressed. 
                         Obviously speculative.

                    - poor research doesn't help - one of the most quoted studies of the
                    cheetah (a book by Eaton) is based on about three months' field
                    research.

               There are cheetahs in zoos, but due to the lack of breeding success zoos
               are reluctant to "import" more.  Particularly as wild populations are also
               declining. 

               Long term outlook is not promising.

          Genetic issues:

               Several studies from the early 1980's indicate that the cheetah lacks
               genetic diversity:

                    - Some speculation that most cheetahs (at least in Africa) are
                    derived from a population bottleneck that occurred about 10,000
                    years ago.

                    - The contribution of this information to conservation is not clear. 
                    It is obvious, for instance that cheetahs were doing quite well even
                    with their low genetic variability until humans started hunting and
                    encroaching on habitat.

                    - Nevertheless, it might help explain the poor breeding success in
                    zoos (though again, they seem to manage fine in the wild).

                    - It does indicate that cheetahs may be more vulnerable to specific
                    threats such as diseases since variability is so low.

               Bottom line - the low genetic variability is interesting and should be kept
               in mind, but it's impact on most conservation issues in probably not that
               important.

          Another warning about web-based research:

               - One web site mentioned:

                    "In Namibia, where the largest populations of cheetahs are found,
                    more than 95% live on private ranchlands".

               - This seems reasonable.  But then another web site seems to have
               interpreted this as:

                    "Currently 95% of all remaining cheetahs live on farmlands in
                    Namibia".

                    (Totally false - Namibia does appear to have the largest remaining
                    population of cheetahs, but East Africa is close behind.  To be
                    honest, numbers are hard to come by, but this is obviously wrong.)

               - It is always best to go to the original sources (journals, papers, etc.).

     That's it for case studies (for a while - we may do more later to illustrate specific points).

     Next we'll spend some time making sure we understand the basic concepts needed to
     make informed decisions about conservation biology.